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The bad wheel Patrick Willis has been playing on for several years finally forced the All-Pro linebacker to make an extended pit stop. Willis was placed on season-ending injured reserve Tuesday because of a problematic left big toe he injured in a win against the Rams on Oct. Willis’ issues with his toe date back several years, he said, and he likened playing with the injury to driving on bald tires. Willis was hopeful he could return this season, but he decided to undergo surgery because he feared the injury could shorten a career that’s included a Pro Bowl berth in each of his first seven seasons in the league.
The loss of Willis is a significant blow for a defense that’s still without All-Pro inside linebacker NaVorro Bowman, but it’s softened somewhat by the presence of rookie linebacker Chris Borland. Borland and Michael Wilhoite will start for the fourth straight game when the 49ers visit the Giants on Sunday.


The 49ers hope Bowman will be playing relatively soon after he tore the ACL and MCL in his left knee in the NFC Championship Game in January. On Sunday, Bowman told ESPN he’s still experiencing knee pain and was unsure when he could return to the practice. The only backup inside linebacker on the 49ers’ current roster is Nick Moody, a 2013 sixth-round pick who has played 10 career defensive snaps, all last year. Meanwhile, Willis is already looking ahead to 2015, when he hopes to take the field with his left foot at full strength for the first time in several years.
In his past two starts subbing for Willis, Borland has collected 35 tackles, included five for losses, and added a fumble recovery. Jim Harbaugh said Tuesday on KNBR that Bowman could “possibly” return to practice soon, but that decision won’t be made until the three-time All-Pro has another MRI.


The 49ers addressed the position Tuesday by re-signing rookie Shayne Skov to their practice squad.
Willis, who was initially optimistic he’d return soon, has missed three games and decided to undergo season-ending surgery when his ailment showed no signs of improvement. Skov, who played at Stanford, spent the offseason with the 49ers and began the season on their practice squad.



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